The human race has been taken over by small parasitic aliens called "Souls". The Souls travel to distant planets inserting themselves individually into a host body of that planet's dominant species. They are able to access the host's memories, and occupied hosts are identifiable by silver rings that form in the hosts' eyes.
Melanie Stryder is captured and infused with a soul called "Wanderer." Wanderer is asked by Seeker to access Melanie's memories in order to discover the location of a pocket of unassimilated humans. Surprisingly, Melanie's consciousness has not been eliminated, and it struggles for control of her body. Melanie and Wanderer carry out an internal conversation with each other, forming something of a friendship.
Wanderer shares with Seeker that Melanie was traveling with her brother, Jamie, and her boyfriend, Jared Howe, to find Melanie's uncle Jeb in the desert. Wanderer admits that Melanie is still present, so Seeker decides to be transferred into Melanie's body to get the information herself. With the help of Melanie, Wanderer escapes and makes her way to the desert, where she is found by Jeb, who takes her to a series of caves hidden inside a mountain where the humans (including Jared and Jamie) are hiding.
Wanderer's presence in the shelter is met with hostility by all but Jeb and Jamie. Seeing this, Melanie instructs Wanderer not to tell anyone she is still alive, since it would merely provoke them, though she later allows her to tell Jamie. Wanderer begins interacting with the humans and slowly begins to gain their trust, forming a bond with Ian O'Shea.
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